"Although we cannot identify the specific weapon that caused this injury, it would be consistent with a halberd or similar weapon.
A boar badge - the symbol of Richard's household - was found at Bosworth and may mark his final stand
"This wound is likely to have been fatal, although that would have depended on exactly how far the blade penetrated into the brain."
A second wound on the other side of the spine indicates where a blade was forced deep into the skull.
Another wound had taken a small chunk out of the top of the head.
"This wound was caused by something hitting the top of the skull sufficiently hard to push in two flaps of bone on the inside surface. Although it looks dramatic, this wound would probably not have been fatal," said Dr Appleby.
She added: "We have evidence of significant injuries on the skeleton, but we cannot conclusively prove that they were the cause of death.
"There are many ways of killing someone that leave no traces on the bones, even in a battle situation."
Robert Woosnam-Savage is curator of European edged weapons at the Royal Armouries Museum in Leeds and has also studied Richard's skeleton.
He said: "Medieval battlefields saw an array of weapons used, from swords, battle hammers, maces, arrows and even early firearms.
"Most of the 'common' foot soldiers would have used staff weapons, such as bills or halberds, which have heavy cutting blades, often with a spike, mounted on a long wooden haft.
"Many of these were designed to be able to punch through elements of armour, or at least damage it in such a way that it no longer functioned."
Mr Woosnam-Savage said using the physical evidence and historical accounts, a possible scenario could be imagined.
Halberds were among the most common weapons used by foot soldiers at Bosworth
"Richard probably got within a few yards of Henry before his horse possibly became stuck in marshy ground or was killed. On foot, with foot soldiers closing in, the fight becomes a close infantry melee.
"It would have been difficult to get through the armour, so attackers would have gone for gaps, or tried to break pieces off.
"The skeleton only shows the minimum number of injuries - the soft tissue has gone - and he is likely to have taken many more wounds of which there is now no trace.
"At some point he loses his helmet and then the violent blows start raining down on the head, including a possible blow from a weapon like a halberd, including the one which I think kills him.
"Then I think it possible that someone has come along, almost immediately afterwards, possibly with his body lying face down and stuck a dagger into his head.
"From becoming unhorsed, it probably only took a matter of a few minutes, before he was dead - not a long time at all."
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